Alison Dos Santos ended Karsten Warholm's reign as hurdles king at the world championships on Tuesday while Britain's Jake Wightman won his country's first 1500m gold in 39 years on a day of upsets. Warholm may have been behind one of the most iconic moments in Olympic history when he smashed the 29-year-old world record to win the 400m hurdles at the Tokyo Games in a time of 45.94 sec. But the 26-year-old came to Eugene on the back of a hamstring injury which ultimately put paid to his medal attempt here. Instead, Dos Santos ran the third fastest time of all time and a championship record of 46.29 sec to win gold ahead of Americans Rai Benjamin and Trevor Bassitt.